(1) Amplitude in the variation of recruitment, survivorship and fecundity was examined for the introduced annual grass Bromus tectorum in three habitat types in eastern Washington (U.S.A.) for three consecutive generations. A total of 18 143 individuals in populations varying from 364 to 5322 members per site were mapped repeatedly from emergence to death with sufficient frequency to detect multiple constituent cohorts varying in age from fewer than 16 to more than 200 days. (2) Recruitment was usually concentrated in late summer and autumn, but occurred at any time until mid-May of the following year. (3) Most of any population experienced low death risk until June, although some cohorts emerging in late summer were devastated (Deevey [...]