Seedlings of the succulent crassulacean acid metabolism (CAM plant Agave deserti in the northwestern Sonoran Desert were found only in sheltered microhabitats, nearly all occurring under the canopy of a desert bunchgrass, Hilaria rigida. Apparently because soil surface temperatures can reach 71@?C in exposed areas, seedlings were generally located near the center or on the northern side of this nurse plant. Both species have shallow root systems, about half of the roots of H. rigida and all those for seedlings of A. deserti occurring above soil depths of 0.08 m. To examine competition for water between the nurse plant and an associated seedling, a three-dimensional model for root water uptake was developed. The model divided the soil [...]