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The restoration of historical fire regimes is often a primary objective in the conservation of fire-adapted forests. However, individual species’ responses to future climate change may uncouple historical vegetation–disturbance relationships, producing potentially negative ecological consequences to fire restoration. We used a landscape simulation model to assess how forest pattern will respond to future climate regimes and whether the restoration of historical fire regimes will benefit forest conservation under future climate regimes. Our study landscape was the 335,000-ha Kaibab Plateau at the North Rim of the Grand Canyon spanning a broad elevation-vegetation gradient of pinyon-juniper, ponderosa pine, mixed...