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We studied the associations of crayfish with reach-scale instream habitat and water quality variables and watershed-scale variables of riparian and watershed land cover, runoff, impervious surfaces, and hydrologic soil type in west-central Indiana. Crayfish assemblage was measured by abundance, species richness, and diversity (Shannon's H) at 180 sites. The western portion of the study area, within the Interior River Lowland ecoregion, has been heavily impacted by agriculture, while the eastern portion, within the Interior Plateau ecoregion, is more heavily wooded and less impacted. The ability of variables at each spatial extent to predict crayfish assemblage was assessed using multiple linear regression analysis....