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Floodplain plant-herbivore-hydroperiod interactions have received little attention despite their potential as determinants of floodplain structure and functioning. We used five types of exclosures to differentially exclude small-, medium-, and large-sized mammals from accessing Fremont cottonwood (Populus deltoides Marshall subsp. wizlizenii (Watson) Eckenwalder) seedlings and saplings growing naturally on four landform types at an alluvial reach on each of two rivers, the Green and Yampa, in Colorado and Utah. The two study reaches differed primarily as a result of flow regulation on the Green River, which began in 1962. Landforms were a rarely flooded portion of the alluvial plain, geomorphically active slow-...