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Geographically Isolated Wetlands (GIWs) occur along gradients of hydrologic and ecological connectivity and isolation, even within wetland types (e.g., forested, emergent marshes) and functional classes (e.g., ephemeral systems, permanent systems, etc.). Within a given watershed, the relative positions of wetlands and open-waters along these gradients influence the type and magnitude of their chemical, physical, and biological effects on downgradient waters. In addition, the ways in which GIWs connect to the broader hydrological landscape, and the effects of such connectivity on downgradient waters, depends largely upon climate, geology, and relief, the heterogeneity of which expands with increasing scale. Developing...