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Conclusions: Assessments of metapopulation structure must consider landscape pattern, but also the non-linear responses of organisms to such patterns Thresholds/Learnings: Synopsis: This study used beetles to empirically test the reliability of neutral percolation models to predict critical thresholds of landscape connectivity. Beetle movements declined sharply when grass cover dropped below 20% of the experimental plot. The findings of this study differed from what was predicted by the model, indicating that landscape connectivity is not dependent on spatial pattern alone, but is also highly dependent how individual organisms move within and among patches depending on the amount of cover. The results suggest that...