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Insect outbreaks are significant disturbances in forests of the western United States, with infestation comparable in area to fire. Outbreaks of mountain pine beetle (Dendroctonus ponderosae Hopkins) require life cycles of one year with synchronous emergence of adults from host trees at an appropriate time of year (termed “adaptive seasonality�) to overwhelm tree defenses. The annual course of temperature plays a major role in governing life stage development and imposing synchrony on mountain pine beetle populations. Here we apply a process-based model of adaptive seasonality across the western United States using gridded daily temperatures from the Vegetation/Ecosystem Modeling and Analysis Project (VEMAP)...