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Conserving a network of representative physical environments is a strategy that could conserve species both now and into the future, while allowing them to move in response to the climate. The key to implementing such a strategy is to define the physical environments in a way that is maximally relevant to species. We created comprehensive maps of geology, elevation zones, and landforms for the Eastern U.S. and then examined the relationships between these factors and 3434 rare species at 85,613 locations. Based on an index of importance derived from the number of species samples and an index of the percent above expected that a species was found on a feature, most species showed strong preferences for specific settings:...