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The desert tortoise Gopherus agassizii is federally listed as Threatened because of documented declines in some populations, yet the proximate causes of these declines are not well understood. With use of radiotelemetry, I monitored a total of 55 individual tortoises at two Mojave Desert sites over three years. Both populations suffered high adult mortality during an extreme drought period, but the temporal pattern and inferred proximate causes of mortality differed between sites. At the eastern Mojave site, no telemetered tortoises died in 1988 or 1989, but 41% died in 1990. All nine carcasses were found and only one showed any evidence of predation or scavenging. Tortoises that died had symptoms of dehydration...